Bothremydidae

Taxonomy

The family is split into two subfamilies and a number of tribes.

Bothremydidae

• Pleurochayah appalachius Adrian et al., 2021 Woodbine Formation, Texas, Late Cretaceous (Cenomanian)
• "Chrysemys" montolivensis Roman 1897 Montoulieu, France, Oligocene (Rupelian)
• Sindhochelys ragei Lapparent de broin et al., 2021 Khadro Formation, Pakistan, Paleocene (Danian)
• Apertotemporalis Stromer, 1934 Bahariya Formation, Egypt, Late Cretaceous (Cenomanian)
• Subfamily Bothremydinae Gaffney et al., 2006
  • Abalakemys Pérez-García, 2023 Farin-Doutchi Formation, Niger, Late Cretaceous (Maastrichtian)
  • Tribe Bothremydini Gaffney et al., 2006
  • Palauchelys López-Conde et al. 2021 Olmos Formation, Mexico, Late Cretaceous (Campanian)
  • Khargachelys AbdelGawad et al. 2021 Quseir Formation, Egypt, Late Cretaceous (Campanian)
    • Subtribe Bothremydina Gaffney et al., 2006
      • Akoranemys Pérez-García, 2021. Madagascar, Late Cretaceous (Cenomanian)
      • Algorachelus Pérez-García 2017
        • Algorachelus parvus (Haas 1978) Bet-Meir Formation, Amminadav Formation, Palestine, Late Cretaceous (Cenomanian)
        • Algorachelus peregrinus Pérez-García 2017 Utrillas Formation, Spain, Nazaré, Portugal, Late Cretaceous (Cenomanian)
        • Algorachelus tibert (Joyce et al. 2016) Naturita Formation, Utah, Late Cretaceous (Cenomanian) (alternatively considered the only member of the genus Paiutemys)
      • Araiochelys Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
        • Araiochelys hirayamai Gaffney et al., 2006
      • Bothremys Leidy, 1865 (jr synonym: Karkaemys)
        • Bothremys arabicus (Zalmout et al., 2005) Wadi Umm Ghudran Formation, Jordan, Late Cretaceous (Santonian)
        • Bothremys cooki Leidy, 1865 Merchantville Formation, Navesink Formation, New Jersey, Tar Heel Formation, North Carolina, Late Cretaceous (Campanian-Maastrichtian)
        • Bothremys kellyi Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
        • Bothremys maghrebiana Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
      • Chedighaii Gaffney et al., 2006
        • Chedighaii barberi (Schmidt, 1940) Brownstone Marl, Arkansas, Blufftown Formation, Georgia, Marshalltown Formation, New Jersey, Tar Heel Formation, North Carolina, Late Cretaceous (Campanian)
        • Chedighaii hutchisoni Gaffney et al., 2006 Cerro del Pueblo Formation, Mexico, Late Cretaceous (Campanian) Kirtland Formation, New Mexico, Tar Heel Formation, North Carolina, Late Cretaceous (Campanian)
      • Rosasia Costa, 1940 Argilas de Aveiro Formation, Portugal, Late Cretaceous (Campanian-Maastrichtian)
        • Rosasia soutoi Carrington da Costa, 1940
      • Inaechelys Carvalho, 2016 Maria Farinha Formation, Brasil, Paleocene
        • Inaechelys pernambucensis Carvalho, 2016
      • Zolhafah Lapparent de Broin and Werner, 1998 Dakhla Formation, Egypt, Late Cretaceous (Maastrichtian)
        • Zolhafah bella Lapparent de Broin and Werner, 1998
    • Subtribe Foxemydina Gaffney et al., 2006
      • Elochelys Nopsca, 1931 France, Spain, Late Cretaceous (Campanian-Maastrichtian)
        • Elochelys perfecta Nopsca, 1931
      • Foxemys Tong et al., 1998 Csehbánya Formation, Hungary, Late Cretaceous (Santonian) Argiles et Grès à Reptiles Formation, Marnes d'Auzas Formation, Marnes Rouges Inferieures Formation, Lestaillats Marls Formation, France, Late Cretaceous (Campanian-Maastrichtian), Villalba de la Sierra Formation, Spain, Late Cretaceous (Campanian)
        • Foxemys mechinorum Tong et al., 1998
      • Palemys Gray, 1870
        • Palemys bowerbankii (Owen, 1842) London Clay Formation, England, Eocene (Ypresian)
      • Polysternon Portis, 1882 Argiles et Grès à Reptiles Formation, Rognacian Formation, France, Vitoria Formation, Laño, Tremp Formation, Sierra Perenchiza Formation, Spain, Late Cretaceous (Campanian-Maastrichtian)
        • Polysternon provinciale (Matheron, 1869)
      • Puentemys Cadena et al., 2012 Cerrejón Formation, Colombia, Paleocene
        • Puentemys mushaisaensis Cadena et al., 2012
      • Tartaruscola Pérez-García, 2016 Saint-Papoul, France, Eocene (Ypresian)
      • Iberoccitanemys Pérez-García et al. 2012 Vegas de Matute Formation, Villalba de la Sierra Formation, Spain, Campanian Marnes d'Auzas Formation, France, Maastrichtian
        • Iberoccitanemys atlanticum (Lapparent de Broin and Murelaga, 1996 originally Polysternon atlanticum) syn Iberoccitanemys convenarum (Laurent et al., 2002)
    • Tribe Taphrosphyini Gaffney et al., 2006
      • • Ilatardia Pérez-García, 2019 Farin Doutchi Formation, Niger, Late Cretaceous (Maastrichtian)
      • Subtribe Nigermydina Gaffney et al., 2006
        • Arenila Lapparent de Broin and Werner, 1998 Dakhla Formation, Egypt, Late Cretaceous (Maastrichtian)
          • Arenila krebsi Lapparent de Broin and Werner, 1998
      • Subtribe Taphrosphyina Gaffney et al., 2006
        • Azabbaremys Gaffney et al., 2001 Teberemt Formation, Mali, Paleocene
          • Azabbaremys moragjonesi Gaffney et al., 2001
        • Eotaphrosphys Perez-Garcia, 2018 Mont-Aimé, France, Late Cretaceous (Maastrichtian)
          • Eotaphrosphys ambiguum (Gaudry, 1890)
        • Labrostochelys Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
          • Labrostochelys galkini Gaffney et al., 2006
        • Motelomama Perez-Garcia, 2018 Salina Group, Peru, Eocene (Ypresian)
          • Motelomama olssoni (Schmidt, 1931)
        • Phosphatochelys Gaffney and Tong, 2003 Ouled Abdoun Basin, Morocco, Paleocene
          • Phosphatochelys tedfordi Gaffney and Tong, 2003
        • Rhothonemys Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
          • Rhothonemys brinkmani Gaffney et al., 2006
        • Taphrosphys Cope, 1869 (jr synonyms: Amblypeza, Bantuchelys, Prochonias)
          • Taphrosphys congolensis (Dollo, 1912) Landana Formation, Angola, Paleocene
          • Taphrosphys dares Hay, 1908 (nomen dubium)
          • Taphrosphys ippolitoi Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
          • Taphrosphys sulcatus (Leidy, 1856) Hornerstown Formation, New Egypt Formation, New Jersey, Late Cretaceous (Maastrichtian)
        • Ummulisani Gaffney et al., 2006 Morocco, Eocene (Ypresian)
          • Ummulisani rutgersensis Gaffney et al., 2006
        • Crassachelys Bergounioux, 1952 (nomen dubium)
          • Crassachelys neurirregularis Bergounioux, 1952
        • Eusarkia Bergounioux, 1952 (nomen dubium)
          • Eusarkia rotundiformis Bergounioux, 1952
        • Gafsachelys Stefano, 1903 (nomen dubium)
          • Gafsachelys phosphatica Stefano, 1903
          • Gafsachelys moularensis Bergounioux, 1955
  • Tribe Cearachelyini Gaffney et al., 2006
    • Cearachelys Gaffney et al., Santana Formation, Brazil, Early Cretaceous (Albian)
      • Cearachelys placidoi Gaffney et al., 2001
    • Elkanemys Maniel, de la Fuente and Canale, 2021 Candeleros Formation, Argentina, Late Cretaceous (Cenomanian)
      • Elkanemys pritchardi Maniel, de la Fuente and Canale, 2021
    • Itapecuruemys Batista et al., 2020 Itapecuru Formation, Brasil, Early Cretaceous (Aptian)
      • Itapecuruemys amazoniensis Batista et al., 2020
    • Galianemys Gaffney et al., 2002 Kem Kem Group, Morocco, Late Cretaceous (Cenomanian)
      • Galianemys emringeri Gaffney et al., 2002
      • Galianemys whitei Gaffney et al., 2002
• Subfamily Kurmademydinae Gaffney et al., 2006
  • Tribe Kurmademydini Gaffney et al., 2006
    • Jainemys Joyce & Bandyopadhyay, 2020 Lameta Formation, India, Late Cretaceous (Maastrichtian)
      • Jainemys pisdurensis Joyce & Bandyopadhyay, 2020
    • Kinkonychelys Gaffney et al., 2009 Maevarano Formation, Madagascar, Late Cretaceous (Maastrichtian)
      • Kinkonychelys rogersi Gaffney et al., 2009
    • Kurmademys Gaffney et al., 2001 Kallamedu Formation, India, Late Cretaceous (Maastrichtian)
      • Kurmademys kallamedensis Gaffney et al., 2001
    • Sankuchemys Gaffney et al., 2003 Intertrappean Beds, India, Late Cretaceous (Maastrichtian)
      • Sankuchemys sethnai Gaffney et al., 2003