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Chirotherium

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Through Germany, England and North America

The creatures who made the footprints and tracks were probably pseudosuchian archosaurs related to the ancestors of the crocodiles. They likely belonged to either prestosuchidae or rauisuchidae groups, which were both large carnivores with semi-erect gaits.

History

Chirotherium tracks were first found in German Triassic sandstones in 1834, and later in England in 1838. They were found before dinosaurs were known and initial models of the trackmaker proposed that it was a bear or ape, which walked with its feet crossed. This proposal was necessary to explain the toe on the outside. The tracks were also proposed to be from a marsupial. These fossil tracks have now been found on North America, Argentina, North Africa, Europe, and China.

British paleontologist Richard Owen suggested in 1842 that the tracks were made by a labyrinthodont amphibian. Over the following years, new discoveries of archosaurian reptiles indicated that Chirotherium tracks were made by a pseudosuchian. The print's resemblance to mammals was only superficial; in reality, an external (lateral) ‘thumb’ was commonplace among Triassic archosaurs.

In 1965, the skeleton of an animal probably closely related to the trackmaker of Chirotherium was found, called Ticinosuchus. It had the external toe on its hind feet but not on its front feet and was possibly a more derived descendant, whose gait did not require a stabilizing front toe. Footprints of different size and proportions occurring together on one and the same bedding plane probably reflect a sex difference (sexual dimorphism) within the trackmaker species.

Other Chirotheriidae include Isocheirotherium.

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