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Galaxiidae

Family of mostly small freshwater fish

  • 5 min read
  • 3 chapters
A member of the Mountain Galaxias species complex that was caught on a tiny barbless wet fly in an upland stream that was free of exotic Trout at the time of capture. This upland stream has since been
Codman at en.wikipedia · CC BY-SA 3.0

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Through Fish, South America and Temperate climate

LabWho Lives Where?

Some galaxiids live in fresh water all their lives, but many have a partially marine lifecycle. In these cases, larvae are hatched in a river, but are washed downstream to the ocean, later returning to rivers as juveniles to complete their development to full adulthood. This pattern differs from that of salmon, which only return to fresh water to breed, and is described as amphidromous.

Freshwater galaxiid species are threatened, some gravely, by exotic salmonid species, particularly trout species, which prey upon galaxiids and compete with them for food. Exotic salmonids have been introduced by humans to many different land masses (e.g. Australia, New Zealand), with little apparent thought as to ecological impacts on native fish. Numerous localised extinctions of galaxiid species have been caused by the introduction of exotic salmonids, and a number of freshwater galaxiid species are threatened with overall extinction by exotic salmonids.

Evolution

Phylogenetic evidence alternatively places galaxiids within the Protacanthopterygii, or more recently as the sister group to the Neoteleostei. Their ancestors are thought to have diverged from the neoteleosts around the Triassic-Jurassic boundary.

The earliest definitive fossils of galaxiids are from the Miocene of New Zealand, which can be placed in the extant genus Galaxias. This young fossil range contrasts with the presumed ancient origins of the group. In 1998, a possible Late Cretaceous (Maastrichtian) galaxiid from South Africa was described as Stompooria. However, later studies have questioned this assignment, as Stompooria differs from galaxiids in many morphological traits, especially in the presence of scales, although it being an ancestral galaxiid that had not yet developed galaxiid traits could not be ruled out. Other taxonomic treatments have instead placed Stompooria as part of an extinct clade sister to the Esociformes and Salmoniformes.

Many galaxiid genera show an unusual form of sister group relationship to one another, where a genus in Australasia is more closely related to a genus in South America than to other genera from the same continent. An example is the relationship between the South American Aplochiton and the Australian Lovettia. It is thought that the evolution of galaxiids may have been influenced by the Antarctic land bridge between Australia and South America, which served as a shallow marine and fresh water connection that allowed different galaxiid taxa to disperse between both continents.

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