Leptothecata

In the sessile stage, Leptothecata are surrounded by a chitinous outer layer as their exoskeleton, including the gonophores, their reproductive organ. Leptothecata exhibit radial symmetry, and their gonads can be found in the radial canals of the medusa stage. Their habits range from benthic to planktonic. The polyps and colonial forms are benthic, whilst the medusae are planktonic. Leptothecata exhibit extensive, complex variation. Thecata colonies also have extensive specialization due to their polyps' function and variation. Most Leptothecata possess statocysts, which are used for defence and protection. The classes that have lost their statocysts have been changed ancestrally over time rather than a direct loss.

The approximately 1,900 species of Leptothecata are characterized by a number of features: Their polyps are always living in colonies with the hydranths set in hydrotheca which are usually permanent and often long enough so the animal can fully retract into it; some have very reduced hydrothecae resembling Anthoathecata. There is a single whorl of tentacles.

The gonophores are borne on much reduced hydranths and usually protected in a peridermal gonotheca. Medusae forming on fully developed hydranths are extremely rare; usually the gonophores develop into medusae or into sessile sporosacs. The medusae have a shallow bell, bear the gonads on their radial canals, and usually have statocysts which are formed only from epidermal tissue and more than four tentacles and. The cnidome never has stenoteles.

Characteristics

Colony architecture among Leptothecata comprises extensive diversity found in the hydrozoans. Their life cycles have been found to be connected with changes in colony shapes. Zooid polymorphism within the colonies are usually specialized. Polyps that make up the colonies tend to have three specialities and functions. First being the gastrozooid, which has their speciality of nutrition and digesting the food. Second, the gonozooid which is the reproductive polyp. Third, being the dactylozooid which function works in defense for the colony. The dactylozooid recently had become more highly variable with not being present in some thecata colony forms, and only possessing the gastrozooid and gonozooid polyps.

Thecata colonies have detectable shapes and arrangements allowing for distinguishing classification between one another. One major shape is when the colonies are erected off a branched colony. Another major shape of thecata is where the colonies can be erected off an unbranched stem. Stolonal colonies are a final major type where their polyps are connected to the creeping part of the colony. Where most cases of the erected branched shape have been found to be derived over time.

Medusae tend to be pelagic. But there are specific medusae species while at the medusa stage can still remain benthic. Polyps can also be free floating, which are called pelagic polyps. Similarly, their gastroular and nervous system have great complexity, as well as their shape. While medusae due tend to lack any presence of visible sense organs. Leptothecata have significant synapomorphies that are present in most of all their species. In regards to their gastrozooids, Lephtothecata have the theca layer on their polyps. Which has allowed the synapomorphy for Leptothecata to form hydrothecae that is also made of theca, that surrounds the gonozooid.