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Ancodonta

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Through North America, Africa and Kenya

The placement of Ancodonta among the artiodactyls originally based on basic morphological work that they were the sister group to pigs and peccaries, placed in the suborder Suiformes. However molecular work has found this not to be the case, as the closest relatives are the cetaceans. Cetaceans and ancodonts are now classified in the suborder Whippomorpha. This led to some theories that the last common ancestors of hippos and whales was a semiaquatic mammal, but in a 2015 genomic study that compared the genes responsible in aquatic adaptations within Whippomorpha found it to be the result of convergent evolution, as opposed to shared ancestry. This resolves the issue of how hippos and whales developed their aquatic adaptation despite modern hippos appearing much later in the fossil record than whales.

The cladogram below shows the results of the phylogenetic analysis conducted by Orliac et al. (2010).

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