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Asymmetry

Absence of, or a violation of, symmetry

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In organisms

Due to how cells divide in organisms, asymmetry in organisms is fairly usual in at least one dimension, with biological symmetry also being common in at least one dimension.

Louis Pasteur proposed that biological molecules are asymmetric because the cosmic [i.e. physical] forces that preside over their formation are themselves asymmetric. While at his time, and even now, the symmetry of physical processes are highlighted, it is known that there are fundamental physical asymmetries, starting with time.

Asymmetry in biology

Asymmetry is an important and widespread trait, having evolved numerous times in many organisms and at many levels of organization (ranging from individual cells, through organs, to entire body shapes). Benefits of asymmetry sometimes have to do with improved spatial arrangements, such as the left human lung being smaller, and having one fewer lobe than the right lung to make room for the asymmetrical heart. In other examples, division of function between the right and left half may have been beneficial and has driven the asymmetry to become stronger. Such an explanation is usually given for mammalian hand or paw preference (handedness), an asymmetry in skill development. Training the neural pathways in a skill with one hand (or paw) may take less effort than doing the same with both hands.

Nature also provides several examples of handedness in traits that are usually symmetric. The following are examples of animals with obvious left-right asymmetries:

  • Most snails, because of torsion during development, show remarkable asymmetry in the shell and in the internal organs.
  • Male fiddler crabs have one big claw and one small claw.
  • The narwhal's tusk is a left incisor which can grow up to ten feet in length and forms a left-handed helix.
  • Flatfish have evolved to swim with one side upward, and as a result have both eyes on one side of their heads.
  • Several species of owls exhibit asymmetries in the size and positioning of their ears, which is thought to help locate prey.
  • Many animals (ranging from insects to mammals) have asymmetric male genitalia. The evolutionary cause behind this is, in most cases, still a mystery.

As an indicator of unfitness

  • Certain disturbances during the development of the organism, resulting in birth defects.
  • Injuries after cell division that cannot be biologically repaired, such as a lost limb from an accident.

Since birth defects and injuries are likely to indicate poor health of the organism, defects resulting in asymmetry often put an animal at a disadvantage when it comes to finding a mate. For example, a greater degree of facial symmetry is seen as more attractive in humans, especially in the context of mate selection. In general, there is a correlation between symmetry and fitness-related traits such as growth rate, fecundity and survivability for many species. This means that, through sexual selection, individuals with greater symmetry (and therefore fitness) tend to be preferred as mates, as they are more likely to produce healthy offspring.

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