Cud

Rumination

The alimentary canal of ruminants, such as cattle, giraffes, goats, sheep, alpacas, and antelope, are unable to produce the enzymes required to break down the cellulose and hemicellulose of plant matter. Accordingly, these animals rely on a symbiotic relationship with a wide range of microbes, which largely reside in the reticulorumen, and which can synthesize the requisite enzymes. The reticulorumen thus hosts a microbial fermentation which yields products (mainly volatile fatty acids and microbial protein), which the ruminant is able to digest and absorb. This allows the animals to extract nutritional value from cellulose which is usually undigested.

The process of rumination is stimulated by the presence of roughage in the upper part of the reticulorumen. The chest cavity is stretched, forming a vacuum in the gullet that sucks the semi-liquid stomach content into the esophagus. From the esophagus it is taken back to the mouth with retro peristaltic movements. When the stomach content, or the cud, arrives in the mouth of the ruminant, it is pushed against the palate with the tongue to remove excess liquid, the latter is swallowed and the solid material is chewed thoroughly so the cattle can extract the minerals present in the cud brought to the surface during rumination. The function of rumination is that food is physically refined to expose more surface area for bacteria working in the reticulorumen, as well as stimulation of saliva secretion to buffer the rumen pH.

When food has been degraded sufficiently it passes from the reticulorumen through the reticulomasal orifice to the omasum followed by the abomasum to continue the digestion process in the lower parts of the alimentary canal. No enzymes are secreted in the rumen. Enzymes and hydrochloric acid are only secreted from the abomasum (fourth stomach) onwards, and ruminants function from that point onwards much like monogastric animals, such as pigs and humans.