Horneophytopsida
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Through Vascular plant
In 2004, Crane et al. published a cladogram for the polysporangiophytes in which the Horneophytopsida are shown as the sister group of all other polysporangiophytes. One other former rhyniophyte, Aglaophyton, is also placed outside the tracheophyte clade, as it did not possess true vascular tissue (in particular did not have tracheids), although its conducting tissue is more complex than that of the Horneophytopsida.
Phylogeny
Partial cladogram by Crane, Herendeen & Friis 2004, with emphasis on horneophytes.
(See the Polysporangiophyte article for the expanded cladogram.)
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Genera
Genera that have been placed in the horneophytes include:
- †Caia Fanning, Edwards & Richardson (1990)
- †Horneophyton Barghoorn & Darrah (1938) (Hornea Kidston & Lang (1920) non Baker (1877); Langiophyton Remy & Hass (1991) form taxon – female gametophyte)
- †Tortilicaulis Edwards (1979)
Other sources place only Horneophyton in this group, treating Caia and Tortilicaulis as actual or possible rhyniophytes.