Pezizales
Order of fungi
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Through Species, Temperate climate and Phylogenetics
Description
Members of this order are characterized by asci that typically open by rupturing to form a terminal or eccentric lid or operculum. The ascomata are apothecia or are closed structures of various forms derived from apothecia. Apothecia range in size from less than a millimeter to approximately 15 centimetres (6 inches), and may be stalked or sessile. The order includes epigeous, semihypogeous to hypogeous (truffles) taxa. The ascospores are single-celled, bipolar symmetrical, and usually bilaterally symmetrical, ranging from roughly spherical to ellipsoidal to occasionally fusoid. The ascospores of some species develop surface ornamentations such as warts, ridges, or spines. The tissues of the ascomata are fleshy and often fragile. Although the majority of species are known only in the teleomorphic state, the anamorphs of some species are known.
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Taxonomy
A phylogenetic analysis of molecular and morphological data suggests that there are three lineages within Pezizales. Lineage A includes Ascobolaceae and Pezizaceae. The former has no known hypogeous species. Lineage B includes the two epigeous families Rhizinaceae and Caloscyphaceae, as well as two sub-lineages with known hypogeous species, Morchellaceae–Discinaceae and Helvellaceae–Tuberaceae. Lineage C include Ascodesmidaceae, Sarcoscyphaceae, Sarcosomataceae with no known hypogeous species as well as Glaziellaceae and Pyronemataceae. It is unknown where Carbomycetaceae fits into this lineage.