Rhinantheae
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The phylogeny of the genera of Rhinantheae has been explored using DNA markers. Three assemblages can be distinguished in this tribe:
- Rhinanthus is the sister genus to Lathraea, and then to Rhynchocorys. These taxa are closely related to the core Rhinanteae.
- In the core Rhinantheae, Odontites sensu lato, including Bornmuellerantha and Bartsiella, is the sister genus to Bellardia, including Parentucellia and Bartsia canescens + B. mutica. These taxa are closely related to Hedbergia (including Bartsia decurva + B. longiflora) and Tozzia. In turn, these genera share phylogenetic affinities with Euphrasia, and then with Bartsia sensu stricto (Bartsia alpina).
- Melampyrum occupies an isolated, deep-branching position.
The median crown age of Rhinantheae was estimated to be ca. 30 Myr.
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Systematics
Rhinantheae is defined as the least inclusive crown clade that includes Pterygiella nigrescens, Rhinanthus cristagalli, Melampyrum pratense, and Tozzia alpina. It comprises 19 genera.
- Bartsia
- Bartsiella
- Bellardia
- Bornmuellerantha
- Euphrasia
- Hedbergia
- Lathraea
- Macrosyringion
- Melampyrum
- Nothobartsia
- Odontitella
- Odontites
- Parentucellia
- Pseuodbartsia
- Pterygiella
- Rhinanthus
- Rhynchocorys
- Tozzia
- Xizangia
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